Senior Research Scientist, Harvard University Herbaria. Specialty: Vascular plants in temperate and subtropical Asia, including the flora of the Transverse Mountains in southwestern China, known as a biodiversity hotspot. He has a deep knowledge of Japanese plants, and has been conducting plant surveys in East Asia since 1977, while Flora of Japan (1992-2020), Flora of China (1989-), Flora of Korea (2001-), He is also a member of the publishing editorial board for many other publications.